The petiole

A leaf (plural leaves) is the major lateral appendage of a vascular plant stem that is normally carried above ground and adapted for photosynthesis. The leaves, stems, blossom, and fruit make up the shoot system. “Autumn foliage,” for example, is the collective name for leaves.

Leaves are available in a wide range of forms, sizes, and textures. Megaphylls are large, flat leaves with complex venation seen on flowering plants, and the majority of species that bore them are known as broad-leaved or Megaphyllous plants.

The plant’s kitchen is referred to as a leaf. This is because they are the principal organ responsible for photosynthesis, which is the process through which the plant acquires its energy or food. Their green tint is due to the presence of chlorophyll.

What is petiole?

The petiole, also known as the mesopodium, is a smooth or grooved cylindrical or subcylindrical stalk of the leaves.

• The petiole raises the lamina above the stem’s level.
• Petiolate leaves have a petiole, whereas non-petiolate leaves or sessile leaves do not have a petiole.
• The petiole aids in the retention of the lamina to the light.
• In the wind, the petiole permits the leaves to flatten.

Leaf Components

The main sections of a leaf are usually the leaf base, petiole, and lamina.

• The base of the leaf is where it connects to the stem. Stipules are two little leaf-like structures at the base of the leaf. This leaf base is wide and hides the stem in monocotyledons like rice, wheat, and other monocotyledons.
• The long, thin stalk that connects the leaf blade to the stem is known as the petiole.
• The leaf blade is another name for lamina. The green, smooth surface of the leaves is what it is. It is made up of veinlets and a short branching vein. The midrib is the vein that goes through the middle of the lamina. The midrib separates the lamina’s surface into two sections. The leaf blade’s veins and veinlets provide stiffness and aid in the movement of water and other chemicals.

Winged petiole

It is the one in which there is a thin flange of tissue along the length of the petiole. It is a green and flattened petiole. Plants like Citrus and Dionaea have winged petiole in them. This also gives a characteristic foliage arrangement to the plant. Outgrowth present on each side of the petiole in some species is called stipules. Leaves that do not have petiole are called sessile or apetiolate. 

1. Plants with compound leaves have their leaflets attached to a continuation in the petiole. This is called the rachis. Each leaflet may be attached to the rachis by a short stalk which is called a petiole.
2. Swollen regions may be present at either end of the petiole and this is known as pulvinar. These are composed of flexible tissue and allow leaf movement. Pulvinar is commonly seen in the bean family- Fabaceae and the prayer plant family- Marantaceae.
3. In some plants, the petioles are flattened and widened to become phyllodes or cladophylls, and here the true leaves may be reduced or absent. Thus, the phyllode itself serves the functions of the leaf. Phyllodes are commonly seen in the genus Acacia, especially the Australian species.

Leaves Modification

We already know that leaves are specialised for photosynthesis. They also have other important responsibilities to play, such as support, food storage, defence, and so on. They have been adjusted in various ways for each of these functions.

Pea tendrils, cactus spines, onion bulbs, insectivorous plant leaves, and other modified leaves are examples. Let’s take a closer look at some of the leaf modifications:

Storage Leaves

Xerophytic plants, such as those in the Crassulaceae family, have thick, succulent leaves that retain water in their tissues. Large vacuoles filled with hydrophilic colloids can be found in the parenchymatous cells of these leaves. This alteration aids the plant’s resistance to desiccation.

Tendrils of the leaves

Plants with weak stems have leaf tendrils. Tendrils are thread-like structures that develop from the leaves. These tendrils sustain the plant by climbing a neighbouring stick or wall. In Lathyrus aphaca, for example, the entire leaf is transformed into tendrils. Pisum sativum’s top leaflets are transformed into tendrils.

Spines of the leaves

Spines are needle-like features that have been adapted into the leaves of a few plants. The spines serve as defence mechanisms. They also cut down on water loss from perspiration. The leaves of Opuntia, for example, are transformed into spines.

Scale Leaves

These are thin, membrane structures with no stalks that seem brownish or colourless. They guard the auxiliary bud that grows in their axil. Onion scale leaves are meaty and thick, and they store both food and water. Sale leaves can also be found in Casuarina and Asparagus.

Hooks for leaflets

The terminal leaflets of some plants are transformed into hook-like features that aid in climbing. Bignonia unguiscati, for example.

Roots of the leaves

One of the leaves present at the nodes is transformed into adventitious roots in a few plants, allowing them to float above the water surface. Salvinia, for example.

Phyllode

The petiole of some plants flattens out and takes the shape of a leaf, turning green in colour. Phyllode is the term for this. Take, for example, Australian Acacia.

Insectivorous Leaves

Only a few plants require nitrogen to grow. The leaves of these plants have been engineered to trap and digest insects. The following are a few of the changes:

• Leaf Pitcher- The leaf lamina of a few plants, such as Nepenthes, is transformed into a pitcher-like structure. The insect is digested by the pitcher’s inner walls, which release a digestive fluid into the cavity.

• Leaf Bladder- In these plants, the leaf segments are transformed into bladders. These plants can be found in bodies of water. Digestive glands are located on the inner wall, which aids in the digestion of the trapped bug. Utricularia, for example.

• In Drosera, the lamina has many hairs at the tip of which is a sticky globule containing digesting enzymes. When an insect lands on the lamina, the hair envelops it.

Leaves’ Functions

The functions of the leaves are as follows:

Photosynthesis

The major function of leaves is photosynthesis. Photosynthesis is the process by which they transform carbon dioxide, water, and UV light into glucose.

Transpiration

The removal of surplus water from plants into the atmosphere is known as transpiration. The opening of stomata in the leaves causes this to happen.

Guttation

Guttation is the process of removing surplus water from the xylem at the edges of the leaves when the stomata are closed.

Storage: Photosynthesis takes place on the leaves. As a result, they conserve water and nutrients. The succulent, thick leaves are especially well-suited to water storage.

Defence

To prevent them from being harmed or devoured by animals, some leaves have been converted into spines. Opuntia, for example.

Conclusion

Petiole connects the lamina to the stem and contains vascular tissues like xylem and phloem that allow the sap to flow through them. Thus, they provide a passage to plant sap, food, and nutrients to the leaf. Photosynthesis takes place in leaves so the petiole is a part of the leaf photosynthesis and produces the products of photosynthesis. Petiole lifts the lamina above the level of the stem. Leaves having petiole are called petiolate leaves and the leaves that do not have petiole are called non-petiolate leaves or sessile leaves. The petiole helps to hold the lamina to the light. Petiole allows the leaves to flatter in the wind.